Interested Article - Водобродка

Водобродка ( лат. Hydraena ) — род жуков-водобродок из подсемейства Hydraeninae . Встречаются повсеместно. Крупнейший род семейства, в котором известно около 1000 видов.

Распространение

Встречаются повсеместно . Для СССР указывалось не менее 12 видов, на Дальнем Востоке России 2 вида ( и ) . Крупнейшая региональная фауна отмечена в Папуа — Новой Гвинее (132 вида) и на Мадагаскаре (106 видов) .

Описание

Водобродки мелкого размера (1—3 мм), удлинённой формы. Усики состоят из девяти-одиннадцати члеников со слабо выраженной четырех- или пятичлениковой булавой. Щупики (ротовые щупальца) намного длиннее усиков. Взрослые жуки растительноядные, личинки плотоядные.

Род Hydraena хорошо определяется следующими аутапоморфными характеристиками: 1) наличие лабрально-мандибулярного сцепления, 2) ментум с острым срединным выступом, 3) различные сложные сенсиллы, и 4) специализированные особенности экзокринной выделительной системы (ESDS). Современные специалисты в целом согласны с тем, что Hydraena является монофилетичным .

Голова сверху мелкопунктированная. Тазики средних и задних ног раздвинутые. В очень длинных нижнечелюстных щупиках 4 сегмента, нижнегубные щупики 3-члениковые. Лапки пятичлениковые, но иногда кажутся 3- или 4-члениковыми из их редукции. Встречаются в текучих, а также стоячих водоёмах (в озёрах и долинах рек) .

Hydraena alticola
Hydraena wolfi
Hydraena jaegeri
Hydraena cooperi
A) H. biltoni ; B) H. minutissima ; C) H. morio ; D) H. nigrita ; E) H. subintegra ; F) H. subintegra ; шкала = 1 мм.

Классификация

Крупный род жуков, известно около 1000 видов но многие виды остаются неописанными. Род Hydraena был выделен в 1794 году немецким энтомологом Johann Gottlieb Kugelann (1753—1815) для вида Kugelann, 1794 .

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- = Shatrovskiy, 1989
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Jäch, 1988 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2014 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1992 ^{i c g}
Jäch & Díaz, 2016
Perkins, 2014 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2014 ^{i c g}
Perkins, 2011 ^{i c g}
Makhan, 2008 ^{i c g}
Perkins, 2014 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 1980 ^{i c g}
Freitag and Jäch, 2007 ^{i c g}
Hatch, 1965 ^{i c g}
Germar, 1824 ^{i c g}
Jäch, 1995 ^{i c g}
(Jäch, 1988) ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1992 ^{i c g}
Perkins, 2014 ^{i c g}
Satô, 1978 ^{i c g}
Perkins, 2011 ^{i c g}
Sainte-Claire Deville, 1905 ^{i c g}
Skale and Jäch, 2011 ^{i c g}
Jäch, 1994 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 1980 ^{i c g}
(Audisio and De Biase, 1995) ^{i c g}
Perkins, 2011 ^{i c g}
Jäch and Díaz, 1999 ^{i c g}
Jäch, 1988 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch and Díaz, 2012 ^{i c g}
Jäch and Díaz, 1998 ^{i c g}
Perkins, 1980 ^{i c g}
Delgado and Palma, 1997 ^{i c g}
Breit, 1916 ^{i c g}
Jäch, 1986 ^{i c g}
Jäch, Díaz and Dia in Jäch, Dia and Díaz, 2006 ^{i c g}
Perkins, 2011 ^{i c g}
Janssens, 1961 ^{i c g}
Perkins, 2011 ^{i c g}
Janssens, 1974 ^{i c g}
Perkins, 1980 ^{i c g}
Apfelbeck, 1909 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 1980 ^{i c g}
Hernando and Ribera, 2001 ^{i c g}
Ganglbauer, 1901 ^{i c g}
Brojer and Jäch, 2011 ^{i c g}
Jäch, 1992 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Freitag and Jäch, 2007 ^{i c g}
Sainte-Claire Deville, 1909 ^{i c g}
Erichson, 1837 ^{i c g}
Jäch, 1992 ^{i c g}
Jäch and Díaz, 2001 ^{i c g}
Jäch, 1992 ^{i c g}
Perkins, 2011 ^{i c g}
Janssens, 1972 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 1980 ^{i c g}
Zwick, 1977 ^{i c g}
Janssens, 1981 ^{i c g}
Jäch and Díaz, 2000 ^{i c g}
Jäch, Díaz and Skale, 2013 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1995 ^{i c g}
Kiesenwetter, 1849 ^b
Kiesenwetter, 1849 ^{i c g}
Perkins, 2011 ^{i c g}
Spangler, 1980 ^{i c g}
Balfour-Browne, 1950 ^{i c g}
Perkins, 2011 ^{i c g}
Janssens, 1981 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 1980 ^{i c g}
Skale and Jäch, 2011 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Orchymont, 1944 ^{i c g}
Orchymont, 1930 ^{i c g}
Orchymont, 1931 ^{i c g}
Jäch and Díaz, 2005 ^{i c g}
Orchymont, 1944 ^{i c g}
Jäch, Díaz and Dia in Jäch, Dia and Díaz, 2006 ^{i c g}
Perkins, 2011 ^{i c g}
Sainte-Claire Deville, 1905 ^{i c g}
Perkins, 2011 ^{i c g}
Brojer and Jäch, 2011 ^{i c g}
Zwick, 1977 ^{i c g}
Perkins, 2011 ^{i c g}
Janssens, 1965 ^{i c g}
Perkins, 2014 ^{i c g}
Kiesenwetter, 1849 ^{i c g}
Orchymont, 1943 ^{i c g}
Jäch, 1988 ^{i c g}
Jäch, 1988 ^{i c g}
Janssens, 1968 ^{i c g}
Orchymont, 1937 ^{i c g}
Rey, 1886 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch and Díaz, 1998 ^{i c g}
Kiesenwetter, 1849 ^{i c g}
Perkins, 1980 ^{i c g}
Janssens, 1963 ^{i c g}
Perkins, 2007 ^{i c g}
Jäch and Díaz, 1998 ^{i c g}
Jäch, 1987 ^{i c g}
Perkins, 1980 ^{i c g}
Chiesa, 1927 ^{i c g}
Perkins, 1980 ^{i c g}
Fauvel, 1903 ^{i c g}
Perkins, 2011 ^{i c g}
Mulsant and Rey, 1852 ^{i c g}
Skale and Jäch, 2011 ^{i c g}
Perkins, 2014 ^{i c g}
Jäch and Díaz, 2006 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1992 ^{i c g}
Skale and Jäch in Jäch and Skale, 2009 ^{i c g}
Freitag and Jäch, 2007 ^{i c g}
Orchymont, 1945 ^{i c g}
Jäch, 1994 ^{i c g}
Perkins, 2007 ^{i c g}
Germar, 1824 ^{i c g}
Perkins, 1980 ^{i c g}
LeConte, 1855 ^{i c g}
Sharp, 1882 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch and Díaz, 2000 ^{i c g}
Waterhouse, 1833 ^{i c g}
(Janssens, 1980) ^{i c g}
Wollaston, 1864 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2007 ^{i c g}
Castro, 2000 ^{i c g}
Lagar, Fresneda and Fernando, 1987 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perez, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2014 ^{i c g}
Zaitzev, 1908 ^{i c g}
Perkins, 2011 ^{i c g}
Zwick, 1977 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 2011 ^{i c g}
Kuwert, 1888 ^{i c g}
(Janssens, 1972) ^{i c g}
Jäch, 1985 ^{i c g}
Jäch, Aguilera and Hernando, 1998 ^{i c g}
Jäch, 1992 ^{i c g}
Orchymont, 1931 ^{i c g}
(Janssens, 1972) ^{i c g}
Kugelann, 1794 ^{i c g}
Guillebeau, 1896 ^{i c g}
Zwick, 1977 ^{i c g}
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Boscá Berga, 1932 ^{i c g}
Curtis, 1830 ^{i c g}
Mulsant, 1844 ^{i c g}
Zwick, 1977 ^{i c g}
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Perkins, 1980 ^{i c g}
Orchymont, 1930 ^{i c g}
Perkins, 2011 ^{i c g}
Orchymont, 1923 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1986 ^{i c g}
Knisch, 1924 ^{i c g}
Brojer and Jäch, 2011 ^{i c g}
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Janssens, 1965 ^{i c g}
Binaghi, 1961 ^{i c g}
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Jäch and Díaz, 1999 ^{i c g}
Orchymont, 1913 ^{i c g}
Orchymont, 1925 ^{i c g}
Orchymont, 1931 ^{i c g}
Skale and Jäch in Jäch and Skale, 2009 ^{i c g}
Jäch, 1988 ^{i c g}
Jäch, 1988 ^{i c g}
Jäch and Díaz, 2001 ^{i c g}
Jäch, 1992 ^{i c g}
Jäch, 1988 ^{i c g}
Jäch and Díaz, 2001 ^{i c g}
Jäch, 1992 ^{i c g}
Ganglbauer, 1901 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 1980 ^{i c g}
Orchymont, 1943 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 1980 ^{i c g}
Janssens, 1974 ^{i c g}
Perkins, 2014 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1992 ^{i c g}
Jäch, 1988 ^{i c g}
Wollaston, 1864 ^{i c g}
Orchymont, 1936 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Skale and Jäch in Jäch and Skale, 2009 ^{i c g}
Rey, 1886 ^{i c g}
Perkins, 2011 ^{i c g}
Kiesenwetter, 1849 ^{i c g}
Jäch, Díaz and Dia in Jäch, Dia and Díaz, 2006 ^{i c g}
Perkins, 1980 ^{i c g}
Orchymont, 1930 ^{i c g}
Orchymont, 1931 ^{i c g}
Blackburn, 1896 ^{i c g}
Zwick, 1977 ^{i c g}
Jäch, Díaz and Skale, 2013 ^{i c g}
Jäch, 1992 ^{i c g}
Sahlberg, 1908 ^{i c g}
Jäch and Díaz, 1999 ^{i c g}
Pretner, 1930 ^{i c g}
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Sharp, 1882 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Orchymont, 1944 ^{i c g}
Baudi, 1882 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 1980 ^{i c g}
Orchymont, 1932 ^{i c g}
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Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 2011 ^{i c g}
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Rey, 1884 ^{i c g}
Jäch, 1988 ^{i c g}
Rey, 1885 ^{i c g}
Orchymont, 1944 ^{i c g}
Orchymont, 1944 ^{i c g}
Orchymont, 1944 ^{i c g}
Ganglbauer, 1901 ^{i c g}
Orchymont, 1944 ^{i c g}
Pic, 1918 ^{i c g}
Orchymont, 1930 ^{i c g}
Orchymont, 1945 ^{i c g}
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Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 2007 ^{i c g}
Perkins, 1980 ^{i c g}
Jäch, 1982 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
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Jäch and Díaz, 2012 ^{i c g}
Jäch, 1982 ^{i c g}
Perkins, 1980 ^{i c g}
Perkins, 1980 ^{i c g}
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Perkins, 1980 ^{i c g}
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Perkins, 2011 ^{i c g}
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Jäch and Díaz, 1998 ^{i c g}
Perkins, 2014 ^{i c g}
Perkins, 2011 ^{i c g}
Jäch, 1994 ^{i c g}
Perkins, 2014 ^{i c g}
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Orchymont, 1923 ^{i c g}
Perkins, 2011 ^{i c g}
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Perkins, 1980 ^{i c g}
Perkins, 2011 ^{i c g}
Perkins, 2011 ^{i c g}
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Janssens, 1963 ^{i c g}
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Jäch, 1987 ^{i c g}
Jäch and Díaz, 1998 ^{i c g}
Jäch and Díaz, 2005 ^{i c g}
Jäch and Satô, 1988 ^{i c g}
Perkins, 2007 ^{i c g}
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Perkins, 2007 ^{i c g}
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Jäch, 1988 ^{i c g}
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Jäch, 1992 ^{i c g}
Perkins, 2011 ^{i c g}
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Jäch and Díaz, 2003 ^{i c g}
Perkins, 1980 ^{i c g}
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Perkins, 1980 ^{i c g}
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Базы данных: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net

Дополнение (2017)

Источник (виды расположены в подродовом порядке):

Дополнение (2022)

Источник:

Дополнение (2023)

Источник:

Примечания

↑ Шатровский А. Г. Сем. Hydraenidae — Водобродки ([род] 1. Hydraena — Водобродка) // Определитель насекомых Дальнего Востока СССР. В 6 т. / под общ. ред. П. А. Лера . — Л. : Наука , 1989. — Т. III. Жесткокрылые, или жуки. Ч. 1. — С. 260—264. — 572 с. — 3150 экз. — ISBN 5-02-025623-4 .
↑ Медведев Л. Н. [род] 4. Hydraena — Водобродка // Определитель насекомых европейской части СССР / под общ. ред. Г. Я. Бей-Биенко . — М. — Л. : Наука, 1965. — Т. II. Жесткокрылые и веерокрылые. — С. 91. — 668 с. — 5700 экз.
от 6 ноября 2022 на Wayback Machine . bugguide.net
Perkins P. D. New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2011. — Vol. 2944. — P. 1—417. — doi : .
↑ Perkins P. D. Hydraenidae of Madagascar (Insecta: Coleoptera) (англ.) // Zootaxa : Журнал. — 2017. — Vol. 4342(1). — P. 1—264. — doi : .
Perkins P. D. New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2011a. — Vol. 3074. — P. 1—198. — doi : .
↑ Philip D. Perkins. Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae). (англ.) // Zootaxa : Журнал. — Auckland , New Zealand : (англ.) ( , 2022. — Vol. 5203, no. 1 . — P. 1—66. — ISSN . — doi : . (3 Nov. 2022)
(англ.) . catalogueoflife.org. Дата обращения: 6 ноября 2023. ,
Kugelann J.G. 1794. Verzeichniss der in einigen Gegenden Preussens bis jetzt entdeckten Käfer-Arten, nebst kurzen Nachrichten von denselben. Neuestes Magazin für die Liebhaber der Entomologie 1(5): 513—582. от 18 октября 2022 на Wayback Machine
↑ Jäch, M. A. & J. A. Díaz. New and little known Palearctic species of the genus Hydraena (s.l.) Kugelann XI. Descriptions of three new species from the Greek island of Euboea, and taxonomic notes on the Hydraena (s.str.) dryops complex (Coleoptera: Hydraenidae). Koleopterologische Rundschau, 86: 61—81. (2016).
Jäch, M. A. & J. A. Delgado. An update of the Hydraena fauna of the Comoro Islands (Indian Ocean), with description of H. (Hydraenopsis) maryae sp. n. from Mayotte (Coleoptera: Hydraenidae). Koleopterologische Rundschau, 86: 91-96. (2016).
Jäch, M. A. & J. A. Delgado. Hydraena Kugelann, 1794 (Coleoptera, Hydraenidae) from the Seychelles, Indian Ocean, with description of a new species. ZooKeys, 623: 75-88. (2016).
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. Catalogue of Life . Дата обращения: 24 апреля 2018. 25 апреля 2018 года.
. Дата обращения: 24 апреля 2018. 26 марта 2020 года.
. BugGuide.net . Дата обращения: 24 апреля 2018. 25 апреля 2018 года.
Perkins P. D. New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2023. — Vol. 5367(1). — P. 1—86. — doi : .

Литература

Jäch M. A., Beutel R., Diaz J. A. & Kodada, J. Subgeneric classification, descriptions of head structures, and world check list of Hydraena Kugelann (Insecta: Coleoptera: Hydraenidae) (англ.) // Annalen des Naturhistorischen Museums in Wien, : Журнал. — 2000. — Vol. 102 B. — P. 177–258.
Ordish R. G. Hydraenidae (Insecta: Coleoptera) (англ.) // Fauna of New Zealand : Журнал. — 1984. — Vol. 6. — doi : .
Perkins P.D. (англ.) // Quaestiones Entomologicae : Журнал. — 1980. — Vol. 16. — P. 3–554.
Perkins P. D. A revision of the Australian species of the water beetle genus Hydraena Kugelann (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2007. — Vol. 1489. — P. 1—207. — doi : .
Perkins P. D. New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2011. — Vol. 2944. — P. 1—417. — doi : .
Perkins P. D. New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2011a. — Vol. 3074. — P. 1—198. — doi : .
Perkins P. D. A revision of the water beetle genus Hydraena Kugelann for southern Africa (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2014. — Vol. 3758(1). — P. 1—92. — doi : .
Perkins P. D. Hydraenidae of Madagascar (Insecta: Coleoptera) (англ.) // Zootaxa : Журнал. — 2017. — Vol. 4342(1). — P. 1—264. — doi : .
Perkins P. D. Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2022. — Vol. 5203(1). — P. 1—66. — doi : .
Perkins P. D. New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae) (англ.) // Zootaxa : Журнал. — 2023. — Vol. 5367(1). — P. 1—86. — doi : .
Trizzino M. et al. A molecular phylogeny of the cosmopolitan hyperdiverse genus Hydraena Kugelann (Coleoptera, Hydraenidae) (англ.) // Systematic Entomology : Журнал. — 2012. — doi : .